“…Major land use changes, such as intensification of agriculture and forestry, often associated with fast population growth, have caused forest loss and fragmentation especially in tropical regions (DeFries et al ., ; Lobovikov et al ., ; Hosonuma et al ., ). In contrast, abandonment of agriculture associated with rural depopulation has occurred most frequently in Europe, where it has been a source of either forest expansion in northern countries or land degradation and shrub encroachment in southern regions (Rey Benayas et al ., ; Azevedo et al ., ). Climate change has been reported as the direct cause of shifts in forest species range in different geographic locations (Walther et al ., ; Peñuelas & Boada, ; Kelly & Goulden, ), and for the increased frequency and intensity of disturbances, such as pathogen outbreaks (Harvell et al ., ; Edburg et al ., ), extreme weather events (Allen et al ., , ; Carnicer et al ., ), and wildfires (Westerling et al ., ), with the consequent widespread forest mortality across continents (Peterman & Bachelet, ; Anderegg et al ., , ; Allen et al ., ).…”