“…However, in denser swards, a strong trade-off has been observed between tiller density and fertility, especially as stand ages (Jungers et al, 2017;Hunter et al, 2020a), so that total densities exceeding roughly 1000 tillers m -2 were detrimental to grain yield. Additionally, there is a marked trend of yield decline as stand ages, leading to a drop in the harvest index as high as 50 % (Culman et al, 2023;Duchene et al, 2023). Different productivity dynamics have been observed in previous studies, with yields peaking the first or the second year of cultivation before dropping down (Jungers et al, 2017;Tautges et al, 2018;Bajgain et al, 2020;Bergquist et al, 2022).…”