“…SBT has provided the basis for numerous insights in bioactive flows, for instance cilia-driven symmetry-breaking flow in vertebrates (Smith, Montenegro-Johnson & Lopes 2019), mucociliary clearance (Smith, Gaffney & Blake 2008) and sperm motility (Gaffney et al 2011), amongst others. Indeed, SBT continues to be an invaluable tool for dynamic fluid-filament interaction simulations (Hall-McNair et al 2019;Schoeller et al 2019;Walker et al 2019) where boundary element methods would prove prohibitively costly.…”