“…Measures of trophic niche, such as gut contents, fatty acid composition and stable isotope ratios, can be considered components of an expanded trophic phenotype (Houle, Govindaraju, & Omholt, 2010). These metrics have been primarily used to understand the average trophic position of consumers (e.g., Jackson, Inger, Parnell, & Bearhop, 2011;Happel et al, 2015;Foley et al, 2017), but there is increasing interest in using these metrics to understand how trophic diversity is partitioned among individuals, populations and species (Bolnick et al, 2011;Price & Guglielmo, 2009). Understanding variability in individual-level trophic diversity (Bolnick et al, 2014;Bolnick, Yang, Fordyce, Davis, & Svanbäck, 2002) could also provide insights into the impacts of resource use on individual fitness.…”