“…The soil of the sand-dune habitat consists mainly of large, granular sand particles that provide relatively low levels of water retention, organic matter content, and nutrients to support plant growth (Chiarello and Joesting 2018, Yong and Warkentin 1966). In comparison, inland coastal plain soils consist of a variety of granular sizes, allowing for relatively greater water retention, organic matter, and nutrient availability (Chiarello and Joesting 2018). The difference in soil moisture and nutrient availability between the 2 habitats may lead to different patterns of biomass allocation, root morphology, and/or petiole and leaf cell size (Sultan and Bazzaz 1993a, b).…”