“…In evolutionary perspective, the mystery of extraversion's origins boils down to the question of why natural selection has maintained variability along this continuum, rather than eliminating variation in favor of fixed genotypes or behavioral phenotypes with the highest net fitness (Lukaszewski, 2013;Lukaszewski & Roney, 2011;Nettle, 2005Nettle, , 2006Nettle, , 2011Verweij et al, 2012). Natural selection can theoretically maintain personality variation via fluctuating (including frequency-dependent) selection pressures that modulate the fitness costs and benefits of different personality trait levels (Buss, 2009;Gangestad & Simpson, 2000;Lewis, 2014;Lukaszewski & Roney, 2011;Nettle, 2006;Penke et al, 2007;Tooby & Cosmides, 1990). This theoretical perspective leads to the expectation that personality variation will often be adaptively patterned in relation to circumstances that predicted optimal trait levels under ancestral conditions.…”