“…The analysis of shape change at a macroevolutionary scale has been stimulated by the development of new methodological approximations for the phylogenetic analysis of geometric morphometric (GM) data (Catalano et al, 2010;Klingenberg and Gidaszewski, 2010;Parins-Fukuchi, 2018;Adams and Collyer, 2019). Studies that include the phylogenetic dimension in GM analyses have been oriented to evaluate a variety of questions including shape/function relationship (Cooke and Terhune, 2015;Borgard et al, 2020), shape covariation (Machado et al, 2018), rates of evolutionary change (Houle et al, 2017;Santos et al, 2019), morphological evolution (Stayton, 2004;Duport Bru et al, 2019;Segura et al, 2021), morphological disparity (Stange et al, 2018;Schaeffer et al, 2019), and phylogenetic inference (Ascarrunz et al, 2019). In particular, the role of changes in developmental timing in generating the shape diversity has been the subject of several recent studies in both extant (Esquerré et al, 2017;Morris et al, 2019;Sherratt et al, 2019) and fossil species (Maiorino et al, 2013;Foth et al, 2016;Prieto-Márquez et al, 2020).…”