“…Research in Beringia has long focused on characterizing paleoenvironments and their evolution at a regional level, relying on proxies that provide information on past environments at a low spatial resolution, such as pollen (Edwards et al, 2000) and megafauna fossils (Guthrie, 1990(Guthrie, , 1968a. A different set of proxies that provide paleoenvironmental information at a higher spatial resolution, such as macrobotanical remains (Zazula, Froese, Elias, Kuzmina, & Mathewes, 2007), insects (Elias, 2001), and sedimentological, pedological, biogeochemical, and genetic analyses of lacustrine and terrestrial sediments (Graham et al, 2016;Kaufman et al, 2016;Kokorowski, Anderson, Sletten, Lozhkin, & Brown, 2008;Reuther, 2013;Vachula et al, 2019;Willerslev et al, 2014) has shown the high degree of environmental heterogeneity in Beringia during the terminal Pleistocene and early Holocene.…”