“…However, had younger stages been used, differentiation of the epimatium might have been possible. Previous workers such as Morvan (1983) have equated the entire region that we studied, from epidermis to the outer boundary of the sclerotesta, with the epimatium. Our previous work with Acmopyle (Mill et al ., 2001) suggests that, in that genus at least, this is unlikely and that only some of this tissue in Acmopyle is derived from the epimatium, the remainder being derived from the ovule.…”