“…The APM solution was seminal to several studies that expanded the original solution to account for bedform geometry, bed thickness, heterogeneities of the porous medium, physicochemical properties of the transported substances, e.g., colloids and metals, surface chemographs, unsteady flow conditions, and groundwater discharge [ Packman et al ., , ; Packman and Brooks , ; Marion et al ., , , ; Zaramella et al ., , ; Cardenas et al ., ; Salehin et al ., ; Boano et al ., , ; Wörman et al ., , ; Cardenas and Wilson , , , ; Sawyer and Cardenas , ]. The effect of flow unsteadiness is particularly important as temporal variations of stream discharge are extremely frequent, and the consequent unsteady hyporheic exchange influences subsurface oxygen saturation [ O ' Connor et al ., ], temperature and reactive loads [ Marzadri et al ., , ], nitrogen transformation rates [ Wondzell and Swanson , ; Lautz and Fanelli , ], and mass exchange [ Malcolm et al ., ], thereby controlling the survival of salmonid embryos and the development of microbial biofilms [e.g., Duff and Triska , ; Greig et al ., ; Battin , ; Battin et al ., ; Wood et al ., , and references therein]. Temporal variations of streamflow properties influence the distribution of hydraulic head on the streambed surface.…”