“…However, investigating responses to variability in oceanographic conditions is an important step to predict the consequences of climate on population dynamics of marine species. Climate change impacts marine species mostly indirectly (Sydeman et al., ) by operating primarily via changes in the availability of feeding and breeding habitats (Emmerson & Southwell, ; Fuentes, Limpus, Hamann, & Dawson, ; Hsieh, Kim, Watson, Di Lorenzo, & Sugihara, ; Idjadi & Edmunds, ; Jones, McCormick, Srinivasan, & Eagle, ) and in the productivity and structure of food webs (Carroll et al., ; Edwards & Richardson, ; Forcada, Trathan, Reid, & Murphy, ; Greene, Pershing, Kenney, & Jossi, ; Jenouvrier, Péron, & Weimerskirch, ; Ramírez et al., ; Richardson & Schoeman, ), although direct impacts, such as physiological impacts, have also been reported (Reid et al., ).…”