“…In conifer plantations, the abundance and composition of advance regeneration have been considered good predictors of future regeneration of hardwood species (Nagaike et al, 2003;Ito et al, 2003Ito et al, , 2004Gotmark et al, 2005;Yamagawa et al, 2008;Igarashi and Kiyono, 2008), although these factors vary widely with stand age, previous land-use history, distance from seed sources, fertilisation, topography, geology, and altitude (Guariguata et al, 1995;Halpern and Spies, 1995;Thomas et al, 1999;Ito et al, 2003;Ramovs and Roberts, 2003;Masaki et al, 2004;Utsugi et al, 2006;Sugita et al, 2008;Widenfalk and Weslien, 2009). In older stages of conifer plantations, substantial numbers of hardwood species usually grow to large sizes as advance regeneration Sugita et al, 2008), and thinning does not always enhance species richness (Halpern and Spies, 1995;Wetzel and Burgess, 2001;Widenfalk and Weslien, 2009; but see Moya et al, 2009), likely because advance regeneration can predominate even after thinning.…”