“…Previous studies indicate that some agricultural land use types are frequently used by native species for foraging, breeding, or simply as stepping stones to reach the next habitat patches (Kupfer et al, 2006;Fischer and Lindenmeyer, 2007;Harvey and Villalobos, 2007;Chazdon et al, 2009a;Gilbert-Norton et al, 2010;Vilchez Mendoza et al, 2014). Landscapes that are more heterogeneous, both in composition and configuration, are more likely to include these land use types, and therefore more likely to provide habitat and habitat connectivity for a variety of species than more homogenous landscapes (Daily et al, 2003;Fischer and Lindenmeyer, 2007;Milder et al, 2010;Fahrig et al, 2011). Combined, PE and LE offer a holistic understanding of humanmodified landscapes and link ecology to the social and political implications of environmental change.…”