“…Our knowledge of frugivory patterns in all these taxa is scarce and lists of visitors to three species of the genus Pilosocereus (P. leucocephalus, P. polygonus, and P. purpusii), and a few closely related genera, are the only information available for the entire clade. Three groups of vertebrates, birds, bats, and reptiles are listed in previous studies (Kissling, Böhning-Gaese, & Jetz, 2009;Naranjo, Rengifo, & Soriano, 2003;Ruiz, Santos, Cavelier, & Soriano, 2000;Silva, 1988;Silvius, 1995;Tschapka, Sperr, Caballero-Martínez, & Medellín, 2008;Wendelken & Martin, 1988). Beyond those lists of visitors, Soriano et al (1999) made the only available study that attempts to quantify the relative F I G U R E 1 The Rancho San Ignacio, a private estate near the city of Xalapa, Veracruz, Mexico, has a population of the woolly torch (Pilosocereus leucocephalus) that grows over an ancient basalt lava flow covered with tropical deciduous forest importance of various frugivore visitors to two columnar cacti in Venezuela.…”