“…Catchments consist of different stores of water with a range of ages and specific chemical compositions that contribute to streamflow (Kirchner, 2003). Streamflow during low-flow periods largely originates from groundwater (e.g., Costelloe, Peterson, Halbert, Western, & McDonnell, 2015;Soulsby et al, 2007), including bedrock groundwater (e.g., Oda, Suzuki, Egusa, & Uchiyama, 2013;Sayama, McDonnell, Dhakal, & Sullivan, 2011;Tague & Grant, 2004). The properties of these groundwater stores, and thus their responses to extended dry periods, vary markedly from place to place, so that the magnitude of low flows can also be highly variable in space (e.g., Floriancic et al, 2018;Smakhtin, 2001;Sun, Kasahara, Otsuki, Saito, & Onda, 2017).…”