“…When N . melanostomus migrate to shallow water, they encounter not only many species of piscivorous fishes (Dietrich et al ., ; Reyjol et al ., ; Taraborelli et al ., ), but also water snakes (King et al ., ) and aquatic birds that hunt fishes near the shore (Somers et al ., ; Jakubas, ; Coleman et al ., ), therefore, increasing demands on spatial cognition to locate and remember sheltered locations. Moreover, male N .…”