“…In contrast, stream networks in mountainous catchments are characterized by network expansion and contraction due to different hydrologic periods (e.g., seasonal changes) [e.g., Godsey and Kirchner, 2014;Jencso et al, 2010Jencso et al, , 2009. In such confined mountainous valley settings, subsurface down-valley flow can represent a large portion of persistent flow paths [Castro and Hornberger, 1991;Jackman et al, 1984;Ward et al, 2013b], possibly explaining why order-ofmagnitude changes in stream discharge have been shown to have inconsequential impact on hyporheic exchange [e.g., Ward et al, 2014Ward et al, , 2016Ward et al, , 2017. Groundwater inflow driven by mountainous catchment topology (e.g., hillslope inputs as a function of lateral area), however, can cause spatial fragmentation of the hyporheic flow field [Caruso et al, 2016].…”