“…In the 20 th century, with increasing devastation of natural habitats, and especially disappearance of large marshy areas, the species started to recede rapidly. Climate seems to con-732 A. Bielecki et al (Jab loński 1992a, b;Fritz 1998Fritz , 2000Lenk et al 1999) and records of P. costata (circles) (Augener 1925;Autrum 1932Autrum , 1936Paw lowski 1936;Arndt 1943;Sapkarev 1964Sapkarev , 1975Paw lowski 1968;Sket 1968;Soós 1969;Zinevici 1971;Lukin 1976;Russev & Janeva 1976;Sawyer 1986;Bielecki & Tarnawski 1980). Jab loński 1992b;Zemanek 1992;Najbar 2008) and localities of P. costata (Paw lowski 1968;Dastych 1967;Kufel 1969;Kasprzak 1972;Radkiewicz 1972;Sandner & Wilkialis 1972;Wilkialis 1964Wilkialis , 1970aWilkialis , b, 1971Wilkialis , 1973Agapow 1975Agapow , 1976Agapow , 1977aHajduk et al 1978;Hajduk & Hajduk 1979a, b;Hajduk 1980;Hajduk et al 1980;Najbar 2008) in Poland. tribute additionally to this situation; on the northern distribution border it is one of the factors that seriously limit the population density (M lynarski 1971).…”