The Central and Western Bohemian basins were part of an intramontane depositional area in the Variscan Orogen, in which localised peat-forming wetlands developed in a high-elevation setting (Pešek 1994(Pešek , 2004. The exact elevation remains uncertain but modelling of the palaeogeomorphology and drainage patterns by Opluštil (2005) suggested that they may have been up to 1000 m above sea-level, at least during middle Westphalian times. We will, therefore, refer to them as upland wetlands sensu Thomas & Cleal (2017).Abundant fossil floras in these deposits have attracted palaeobotanists since the early 19 th century (e.g. Sternberg 1820-1838, Ettingshausen 1852, 1854) and are evidently the remains of vegetation broadly similar to that of the lowland paralic swamps of the Variscan foreland (e.g. Cleal 2018) but preserved in a higher-elevation, wetland setting. However, there are clear differences in the species present (Gothan 1915(Gothan , 1951(Gothan , 1954 Cleal 2008a, b) and it is now evident that these upland floras had an important influence on the overall dynamic evolution of the Euramerican peat swamp vegetation (