Winter comprises a substantial fraction of the year across large areas of the globe, but what happens in aquatic ecosystems during winter is not well understood. Recent large-scale observations of decreasing ice cover (Sharma et al., 2016(Sharma et al., , 2019, warming winter temperatures (USGCRP, 2017), and increased recognition of connections between winter dynamics and conditions in other seasons (Cline et al., 2020;Collins et al., 2019;Katz et al., 2015) indicate a need to better understand the fundamental controls on winter conditions and their effects on aquatic processes and communities . We have made recent progress in understanding winter dynamics in lake ecosystems (e.g., Hampton et al., 2017), but we know much less about the winter ecology of rivers even though they are experiencing the same widespread increases in winter temperatures and ice loss (