“…In Equation ( 33), A EHO n,j,z is the newly updated clan and A EHO j,z denotes the previous location of elephant z in clan j, λ 0, 1 ½ is the scaling factor, and it specifies the effect of a matriarch on j on A EHO j,z : A EHO best,j represents matriarch j with the fittest elephant individuals of clan j, r 0, 1 ½ : In each clan, the fittest elephant are not calculated using Equation (33), that is, A EHO j,z ¼ A EHO best,j : Rather, for the fittest elephant, the update process is performed as per Equation (34).…”