“…This is mainly because PA guidelines frequently inhibit some human activities, promoting land abandonment and loss of rurality, with a negative impact on the well-being of local populations, who are vulnerable to the establishment of PAs [1,2,59,81,82]. In recent decades, it is increasingly recognized that PAs must not only focus their conservation efforts on protecting wildlife and landscape naturalness, but also maintaining and supporting the livelihoods of local people by means of more effective policies addressed to protect traditional culture, because most of the PAs of the world show some degree of human use or 'culturalness' [29,83,84].…”