Damselflies of the Family Kennedyidae (Odonata: Archizygoptera) from the Middle–Upper Triassic of Kyrgyzstan

New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata)

“…The Triassic record of stem-relatives of dragon- and damselflies (Odonata), though limited compared to Permian and Jurassic ones, yet indicates that these insects evolved very divergent morphotypes at the time (Deregnaucourt et al 2023). Besides the rare Panodonata, the taxon including crown-Odonata and represented by mid-sized species at the time (Bechly 1997; Nel et al 2002; Pritykina 1981; Tierney et al 2020), the gracile, damselfly-like type was represented by the Archizygoptera (Bechly 1997; Deregnaucourt et al 2021b; Felker 2022; Nel et al 2005; Pritykina 1981). Besides these two taxa, the Triadophlebiomorpha, a group almost exclusively Triassic, included the most large-sized odonatans of the time and evolved a number of peculiar structures, such as a very large area between RP3+4 and MA, a very long petiole, and a number of distinctive vein fusions (Pritykina 1981; Nel et al 2001; Béthoux et al 2009; Béthoux & Beattie 2010; Deregnaucourt et al 2017, 2021a; Zheng et al 2017a, b; Jouault et al 2023).…”