“…The phylogenetic relationships among modern soft-shelled turtle species are still controversial, but it is generally accepted that Trionychidae consists of two clades, Cyclanorbinae and Trionychinae, and that Trionychinae includes some well-supported monophyletic clades (Meylan, 1987;Engstrom et al, 2004). The taxonomy and phylogenetic relationships of fossil trionychid species are far more controversial, and very little is known regarding the origin and early radiation of this group (Gardner et al, 1995;Joyce andLyson, 2010, 2011;Vitek and Danilov, 2010;Vitek, 2012;Danilov and Vitek, 2013;Joyce et al, 2013). The early record of soft-shelled turtles is poor, and most taxa are based either on fragmentary shells or skulls (Yeh, 1994;Hutchison, 2000;Sukhanov, 2000;Danilov and Vitek, 2013).…”