“…The surface holes in the epithelium of the dermal cornea of Geotria australis appear to be a characteristic feature in high abundance with an inverse relationship between the size of the holes and the size of the epithelial cells. The presence of these holes has been previously reported for the Pouched lamprey Geotria australis ( Collin and Collin, 2000a , 2006 ), the Shorthead lamprey, Mordacia mordax ( Collin and Collin, 2006 ), the ammocoete stage ( Dickson et al, 1982 ), but not the adult stage, of the Sea lamprey ( Petromyzon marinus ) ( Van Horn et al, 1969a ; Pederson et al, 1971 ; Dickson et al, 1982 ), the Black shark, Dalatias licha ( Collin and Collin, 2000b , 2006 ), and the Australian lungfish, Neoceratodus forsteri ( Collin and Collin, 2006 ). Surface holes also occur in the pre-metamorphic Axolotl, Ambystoma mexicanum ( Collin and Collin , 2000b , 2006 , 2021a ) but have almost completely disappeared in the post-metamorphic stage ( Collin and Collin, 2021a ).…”