“…The waterway from Chongqing to Yibin in the upper Yangtze River (Figure 1) plays a vital role in the comprehensive transportation system of western developing areas in China (Rohacs & Somongati, 2007), and it is also the foundation for establishing an international shipping center in the upper Yangtze River (Zhao, 2015). Although waterway regulation and dredging projects have significant economic efficiency, their influence on riverine ecosystems cannot be ignored (Cui, Hu, & Zhai, 2011; Nanding et al, 2020; Willems, Busscher, Woltjer, & Arts, 2018). Engineering activities in the river influence the water flow pattern and river connectivity (Grill, Lehner, & Thieme, 2019), which subsequently affects the habitat (Xu, Wang, Duan, & Pan, 2014; Zhou et al, 2019) and species diversity of aquatic organisms (Zhao, Sheng, Yang, Wang, & Zhang, 2015; Zhao, Wang, Pan, Xu, & Li, 2015; Zhou et al, 2017).…”