“…Opitz (, ) erected an additional three subfamilies and attributed subfamily rank to the Isoclerini (of Thanerocleridae), recognized 12 subfamilies (Thaneroclerinae, Isoclerinae, Hydnocerinae, Anthicoclerinae, Tillinae, Clerinae, Epiphloeinae, Tarsosteninae, Peloniinae, Enopliinae, Neorthopleurinae, Korynetinae). Several subfamily‐defining character states, treated as synapomorphies in Opitz' () data matrix, within the taxon descriptions, are qualified with statements such as ‘most genera’ and ‘almost always’, for example, metacoxae with carina for Tillinae and furcal lamina [of metendosternite] absent in Epiphloeinae, whereas another, ‘pronotal commissure partially developed’, is only revealed as synapomorphic because the states of the character ‘extent of development of pronotal commissure’ are scored in a nonlinear way, i.e.…”