“…The projection from medial septum to the dentate gyrus displays one of the more clearly layered patterns of termination by cholinergic a¡erents (Amaral and Witter, 1995;Baratta et al, 1996a,b;Deller, 1998;Deller et al, 1999;Fo « rster et al, 1997;Makuch et al, 2001;Storm-Mathisen and Blackstadt, 1964;Super and Soriano, 1994;Zimmer et al, 1986), and it is this layered quality that makes this region an ideal model system for this study. Septal cholinergic projections to the dentate gyrus terminate principally in the inner molecular layer, although termination also occurs in the outer molecular layer and, to some extent, within the hilus (Amaral and Witter, 1995;Deller et al, 1999;Makuch et al, 2001;Milner et al, 1983;Super and Soriano, 1994).…”