“…Sufficient availability of soil and soil resources at higher elevations (Holtmeier & Broll, ; Butler et al., ; Leonelli et al ., ; Sullivan et al ., ), little dispersal limitation (Holtmeier & Broll, ; Grace et al ., ; Nathan et al ., ; Harsch et al., ), rapid growth (Holtmeier & Broll, ; Lenoir et al ., ; Elmendorf et al ., ,b; Harsch et al., ), short time lags (Brown & Vellend, ), and the absence of confounding interactions with other species (Hille Ris Lambers et al ., ) have been identified in previous studies. Data presented here support the idea that dispersal, plant growth, time lags, and interactions are important, with forests moving upward slowly, perhaps because of competition with tall shrubs.…”