“…Hence, dominant ecological conditions at reefs play an important role in shaping the material record of nearby shell middens. Clearly, there is a need for archaeo-malacology studies, particularly in southern Africa, to engage more fully with ecological variables beyond general or basic considerations in order to maximize data interpretation (e.g., Campbell, 2008;Fa, 2008;Flores and Broitman, 2008;Giovas et al, 2010Giovas et al, , 2013Jerardino et al, 2014;Thakar, 2011). Other important sources of variability in zooarchaeological assemblages, such as human predation, prey choice and foraging strategies, seasonality, and environmental changes, to name a few, are also important and need to be weighed on a case by case basis (e.g., Bird et al, 2002;Giovas et al, 2010Giovas et al, , 2013Jerardino et al, 2008;Whitaker and Byrd, 2012).…”