“…The relatively mild and humid climate on this site allows rapid moraine colonization by plants, causes the accumulation of organic material, and promotes relatively rapid soil development (Alexander and Burt, 1996;Luo et al, 2012). Several studies have investigated how communities develop during primary succession, including pedogenesis (He and Tang, 2008), soil respiration (Luo et al, 2012), plant succession simulation , soil microbial changes , and the availability and transformation of essential nutrients, particularly phosphorus and sulfur speciation (Zhou et al, 2013;Prietzel et al, 2013a, b). However, successional studies have rarely investigated the responses of soil fauna communities to different development stages of ecosystems, and whether these changes parallel those observed in aboveground communities.…”