“…Complex high molecular weight (HMW) polymers are degraded by the soil biota to simple sugars such as glucose, galactose (Gross et al, 1998;Brüll et al, 2000;De Brouwer and Stal, 2001) and fructose, which can be readily utilized as carbon source or energy reserve (Bertocchi et al, 1990) by the crustal microbial community and/or by adjacent plants. Indeed, C from EPSs was demonstrated to be the primary substrate respired in the pulse of CO 2 typically observed in dry lands after rainfall events (Thomas et al, 2008). In other systems it is documented that, owing to microbial activity, EPSs are eventually reduced to smaller polymers with a broad range of molecular sizes (Arnosti,1995(Arnosti, ,1996De Brouwer and Stal, 2001) and chain lengths (Bender et al, 1994), depending on the structure and composition of the microbial community.…”