“…Approximately 340 species have been recognized and grouped into 17 species groups (Lynch, 1976). Evolutionary relationships in the genus are complex because of the overwhelming number of species, the intraspecific variation evident in many species (Cochran and Goin, 1970;Goin, 1947Goin, , 1954Lynch, 1967;Lynch and Duellman, 1980;Rivero, 1978;Schwartz, 1973Schwartz, , 1974Schwartz, , 1976, and the lack of taxonomic characteristics that can be applied to all the species over their entire ranges. The characters used to define species groups in the West Indies are not useful to define groups in South and Central America (Lynch, 1976).…”