“…Similar analyses have been undertaken with aquatic invertebrates (Malmqvist, 2002;Quinlan et al, 2002), flies (Delettre and Morvan, 2000), ants and spiders (Woinarski et al, 2002), saproxylic beetles in woodlands (Ranius, 2002;Similae et al, 2002) and dung beetles (Roslin, 2001). Ground beetles have also been extensively studied because they are easy to catch and are found throughout habitats such as wooded landscapes (Magura et al, 2003;Judas et al, 2002;Koivula and Niemelä, 2002), agricultural land (Kinnunen et al, 2001;Pena et al, 2003), grasslands (Niemelä and Baur, 1998;Vessby et al, 2002) and the urban environment (Andersen, 2000;Niemelä et al, 2002). Much of this research has been based on surveys of individual habitat types, but the diistribution and biodiversity of a number of invertebrate groups agricultural landscapes has also been investigated (Burel et al, 1998;Jeanneret et al, 2003).…”