Modern terrestrial ecosystems on the Earth are characterized by "birds' twitter and fragrance of flowers" (Benton, 2010). The dominant biological groups include flowering plants (angiosperms), insects, mammals and birds (Benton, 2010;Z. Zhou, 2006). These groups closely relate to human society, including food security, disease control and economic development. How the modern terrestrial ecosystems evolved from the ancient ecosystems in the Mesozoic has long been debated (