JSTOR is a not-for-profit service that helps scholars, researchers, and students discover, use, and build upon a wide range of content in a trusted digital archive. We use information technology and tools to increase productivity and facilitate new forms of scholarship. For more information about JSTOR, please contact support@jstor.org.. British Ecological Society is collaborating with JSTOR to digitize, preserve and extend access to Journal of Animal Ecology. Summary 1. Foraging data for great crested grebes (Podiceps cristatus L.) feeding in shallow fish-ponds (0 8-1 m deep) were used to test whether pond characteristics, fish density, mean wet weight of the fish or fish species influenced foraging behaviour and success in this fish-eating grebe. 2. When fishing for roach/rudd (Rutilus rutilus L.IScardinius erythropthalmus L.), the number of prey eaten per unit time foraging was positively influenced by the proportion of successful dives, and negatively by the mean diving time. The proportion of successful dives increased as fish weight decreased. Mean diving time, however, was not related to any component of the food supply, nor to pond characteristics. 3. The overall mean diving time was low in comparison with other studies, probably because the study ponds were so shallow. Since the water depth was equal in each pond, mean diving time is not an accurate measure of fish densities in shallow waters. 4. When roach/rudd were the prey, handling time above the water did not constrain the number of prey eaten per unit time foraging at any fish density. With roach/rudd and carp (Cyprinus carpio L.) as prey, the above-water handling time increased with the size of the fish consumed. The total handling time under and above the water surface together constituted on average 4-7% of the time required to consume one fish. 5. When fishing for roach/rudd, successful dives lasted significantly longer than unsuccessful ones. As the mean duration of successful dives increased with fish weight, we suggest that this can be explained by handling prey under water. For carp, however, this could not be shown. Furthermore, unsuccessful dives lasted significantly longer when fishing for carp than for roach/rudd. This is probably related to behavioural differences between the fish species. In roach/rudd ponds, it seemed that grebes took on average the smaller fish.