“…In this regard, some indications are provided by benthic foraminifera, which are controlled by oxygen and food availability in bottom-waters (e.g., Sliter and Barker, 1972;Nyong and Olsson, 1984;van Morkhoven et al, 1986). In particular, studies on Tethyan sites (Cobianchi et al, 1999;Coccioni et 4 al., 2006;Michalík et al, 2008;Józsa et al, 2016) and sections worldwide (Elkhazri et al, 2013;Bargen and Lehmann, 2014;Zorina et al, 2017) are suggestive of a decrease in benthic foraminifera abundance before OAE 1a. Several locations show scarce or absent benthic specimens during OAE 1a, possibly, in response to a major paleoenvironmental perturbation at the seafloor (Mutterlose and Böckel, 1998;Cobianchi et al, 1999;von Bargen and Lehmann, 2014).…”