“…It is likely that the decline in tiller numbers per marked shoot during the later stages of canopy development was partially due to the competition in the resources between tillers. As the canopy closed, outgrowth of buds from a tiller was reduced by self-shading or shading from the neighboring tillers (Jewiss, 1972;Deregibus et al, 1983;Davies & Evans, 1990;de Kroon & Kwant, 1991;Jonsdottir, 1991;Makita, 1996). As a canopy closes, some old tillers senesce and new tiller production probably becomes a liability rather than a benefit, because the new tillers are being shaded and are less likely to make a positive contribution to the carbon budget of the plant (Skinner & Nelson, 1992).…”