In addition to the large continents of N America, Scandinavia and Gondwana, the regions around the early Palaeozoic Iapetus Ocean contained several small terranes. These included an island arc, which collided with various parts of N America to produce, in succession, the Grampian, Humberian and Taconic orogenies. They also included the terrane of Avalonia, a later Precambrian arc which had rifted off a margin of Gondwana by the middle Ordovician. Large sinistral strike-slip faults in Scotland suggest a total displacement of around 1500 km, so that by the Silurian an elongate Scotland lay to the W of Norway. Continental collisions took place in three stages: a Llandovery stage, perhaps related to eastward subduction below Svalbard, when W-verging nappes were emplaced in E Greenland, a later Silurian (Scandian) stage when westward subduction below Scotland can be related to E-verging nappes in Norway, and an early Devonian stage when Avalonia collided with N America (Acadian orogeny).