The metacarpophalangeal joint is the intermediate joint of the thumb. Its main purpose is to ensure the stability of the grips between the thumb and the fi ngers and the cylindrical grabs using the palm.It allows 60-70° of active fl exion and until 80° of passive fl exion. The amplitude of active and passive extension is 0°.It's a condyloid joint, which means it has 2° of freedom (fl exion/extension and varus/valgus), to which must be added a third one: the automatic axial rotation, essential in the pulp-to-pulp oppositions between the thumb and the fi ngers (with the participation of the trapezio-metacarpal and the interphalangeal joints).This automatic rotation in pronation during the fl exion is related to the combined action of various anatomical elements:• Capsulo -ligamentous , because the ulnar collateral ligament is shorter than the radial collateral ligament. During the fl exion, the ulnar collateral ligament slows the movement of the ulnar part of the fi rst phalanx before the radial collateral ligament holds the radial part back, which creates an automatic pronation of the phalanx on the metacarpal. The same mechanism occurs in extension, leading to a supination of the metacarpophalangeal joint.• Muscular , as the external thenar muscles are more active in fl exion than the internal thenar muscles and provide a combined movement of fl exion-pronationradial inclination.• Osseous , because the internal anterior part of the metacarpal head is higher than the external one. This makes the internal movement of the phalanx shorter than its external movement ( Fig. 6.1 ).