“…It has been shown that most hybrid poplar species are sensitive to O 3 with respect to visible leaf injury (Ryan et al, 2009;Hoshika et al, 2013), damaged photosystems (CO 2 assimilation and chlorophyll fluorescence) (Novak et al, 2005), and reduced growth (Bortier et al, 2000;Hoshika et al, 2013). As the representative deciduous tree species in Northern China, Populus trees are facing the risk of O 3 damage due to high ground-level O 3 pollution in China Yuan et al, 2015).…”