In 2004, a controversy arose in Aotearoa New Zealand's coal mining sector that involved Powelliphanta augusta, a species of land snail. This paper considers that controversy in terms of emergent assemblages involving the snail, the mining industry, the Department of Conservation, political action by environmental non‐government organisations, and public expressions of environmental and other values. Three sets of relations embedded in those assemblages are examined—those related to positioning and organising nature; values, politics, and power relations; and human and non‐human materialities. Those relations illuminate how a protracted controversy and its objects form a “social test bed” for contested views in legislation and guidelines dealing with economy, conservation, and restoration. To assess the dynamics, complexities, and contingencies of such relations, attention is paid to the controversy's detailed time line and legal framing, as well as to ontological shifts of snail and habitat and snail metrics. This analytical focus highlights the shifting quality of ideas about how the snail and its habitat are understood as entangled in fraught and mutating strategies of translocation that are played out in a particular regulatory setting. It also provides insights into international geographical efforts to re‐shape environmental governance and management.