“…Landscapes are generally varied, inhomogeneous, or patchy (e.g., Voorhies, 1972;Fedick and Ford, 1990;Ford and Fedick, 1992;Dunning and Beach, 1994;Fedick, 1995Fedick, , 1996Dunning et al, 1998;Fedick et al, 2000;Kunen, 2001;Penn et al, 2004;Ford et al, 2009;Patterson et al, 2010;Zhang et al, 2010), but logit and WofE models are largely insensitive to landscape variability. This is problematic because settlement behaviour must have been responsive to local conditions while simultaneously determined by larger-scale patterns of landscape use.…”