“…Rich ter res trial flo ras pre served in Eocene sed i ments of this re gion pro vide sig nif i cant clues about Ce no zoic cli mate change. Several au thors have used var i ous bi otic com po nents to re construct depositional and cli ma tic con di tions dur ing the ac cu mu lation of the sed i men tary se quences: wood (Lucas and Lacey, 1981;Fran cis and Poole, 2002), leaves im print (Zastawniak, 1981(Zastawniak, , 1990(Zastawniak, , 1998Zasta wniak et al, 1985;Zastawniak, 1986, 1989a, b;Li, 1992;Dutra, 2001Dutra, , 2004Hunt and Poole, 2003;Kellner et al, 2007;Fontes and Dutra, 2010), plant and fish as sem blage (Doktor et al, 1996), wood, tree-ring and palynomorph set to gether (Askin and Spicer, 1995;Duan and Cao, 1998;Fran cis et al, 2008;Panti et al, 2012). The terres trial flo ras con firmed a warm (i.e., frost-free) Eocene cli mate with a gen eral cool ing trend up the suc ces sion, cul mi nat ing in a sig nif i cant cool ing dur ing the Late Eocene.…”