“…In the real world, fish swimming works in context with many factors, including light and vision (16,24,30), sound (16), social interactions (49), feeding, predators, water quality, chemical cues, fish size and age, and hydraulics at scales smaller than considered here. Differences between modeled and actual environments could have implications on what we infer about fish behavior (50); however, at minimum, our study suggests that abstractions of the real world from hydraulic and behavioral modeling may inform how engineered features function with the cue responses that fish have naturally evolved.…”