Abstract. In the present paper, we study the infinitesimal symmetries of the model of two Riemannian manifolds (M, g) and (M ,ĝ) rolling without twisting or slipping. We show that, under certain genericity hypotheses, the natural bundle projection from the state space Q of the rolling model onto M is a principal bundle if and only ifM has constant sectional curvature. Additionally, we prove that when M andM have different constant sectional curvatures and dimension n ≥ 3, the rolling distribution is never flat, contrary to the two dimensional situation of rolling two spheres of radii in the proportion 1 : 3, which is a well-known system satisfying É. Cartan's flatness condition.