“…Good (1953) first addressed the issue of estimating E (C r ) and proposed an empirical Bayesian approach. Examples of bias due to failing to account for sampling without replacement in species richness estimation can be found in , Haas et al (2006), and Chao and Lin (2012). The Good-Turing method has been applied successfully in several disciplines, such as information retrieval (Song and Croft, 1999), computational linguistics (Church and Hanks, 1990), speech recognition (Jelinek, 1998;Chen and Goodman, 1999), species richness estimation (Esty, 1985;, population size estimation , Shannon entropy estimation , and missile coverage estimation (Lo, 1992).…”