“…This limits plant growth in the later savanna types (Baruch and Go´mez, 1996). Similar overriding influence of nutrient availability on the composition and structure of tropical savannas has been reported in other studies from Venezuela and Colombia (Beard, 1953;San Jose´et al, 1998;Susach, 1989) and from Guyana, Surinam and Brazil (Goodland, 1966;Goodland and Pollard, 1973;Goldsmith, 1974;Lopes and Cox, 1977;Miranda et al, 2003;van Donselaar, 1965). In addition, it has been proposed that fertility edaphic gradients were the main cause for the vegetation gradual transformation from grassland or treeless savanna to the typical woody savanna (Goodland and Pollard, 1973;Lopes and Cox, 1977;Sarmiento and Monasterio, 1969).…”