“…Vannote et al, 1980), and macroinvertebrate distribution was primarily related to the organic energy base, as well as to habitat or biotope preferences (Culp & Davies, 1982;Cushing et al, 1983;Hawkins, 1984;Bott et al, 1983;Wallace, 1988). The role of slope and elevation has been mentioned (Sheldon, 1985;Marchant et al, 1985;Devan & Mudna, 1986;Ormerod & Edwards, 1987;Benke & Meyer, 1988;Gladden & Smock, 1990), but not as explidtly emphasized as, for example, in vegetation ecology, where elevation has long been recognized as a surrogate for a range of environmental gradients which strongly influence vegetation composition (Austin, Cunningham & Good, 1983;Druitt, Enright & Ogden, 1990;Parker, 1991;Palmer & van Staden, 1992). The recognition of variables for which steepness and altitude are surrogates in aquatic ecosystems, is implidt in the long-standing recognition of the 337 importance of factors such as temperature and current speed (Hynes, 1970).…”