Agglutinated Conical Foraminifera (Orbitolinidae, Coskinolinidae) From the Upper Cretaceous (Campanian) of Greece, With Description of Paracoskinolina Klokovaensis N. Sp.

Abstract

An assemblage of agglutinated conical foraminifera including Lepinoconus chiocchinii Cruz-Abad et al., Calveziconus lecalvezae Caus & Cornella, Paleodictyoconus sp., and Paracoskinolina klokovaensis n. sp. are described from the upper lower-middle Campanian of Klokova Mountain of the Gavrovo-Tripolitza Zone, SW continental Greece. With the presence of one rafter in the marginal zone, the new species P. klokovaensis compares to the Lower Cretaceous species Paracoskinolina arcuata (Arnaud-Vanneau) that is distinguished by its cylindroconical test morphology and much larger size. The assemblage occurs in inner platform carbonates associated with other foraminifers such as dicyclinids, and Accordiella conica Farinacci. Representatives of the genus Paracoskinolina were so far only reliably reported from the Upper Berriasian–Albian interval. This new record suggests that the genus either survived the larger benthic foraminifera extinction event associated with the Cenomanian – Turonian boundary anoxic event, or may be an example of an Elvis taxon or homoplasy.